DOI QR코드

DOI QR Code

Cretaceous Pterosaur Tracks in Daegok-ri, Ulju-gun, Ulsan: Spatio-temporal Distribution of Pterosaur in the Korean Peninsula

울산 울주군 대곡리에서 산출된 백악기 익룡 발자국: 한반도 익룡의 시공간적 분포

  • Ha, Sujin (Department of Geological Sciences, Pusan National University) ;
  • Chae, Yong-Un (Department of Geological Sciences, Pusan National University) ;
  • Lim, Jong Deock (Cultural Heritage Administration) ;
  • Kim, Cheong-Bin (Department of Physics Education, Sunchon National University) ;
  • Lim, Hyoun Soo (Department of Geological Sciences, Pusan National University) ;
  • Kim, Kyung Soo (Department of Science Education/Korea Institute of Geoheritage, Chinju National University of Education)
  • 하수진 (부산대학교 지질환경과학과) ;
  • 채용운 (부산대학교 지질환경과학과) ;
  • 임종덕 (문화재청) ;
  • 김정빈 (순천대학교 물리교육과) ;
  • 임현수 (부산대학교 지질환경과학과) ;
  • 김경수 (진주교육대학교 과학교육과/부설한국지질유산연구소)
  • Received : 2018.06.22
  • Accepted : 2018.10.05
  • Published : 2018.10.31

Abstract

Pterosaur tracks were discovered recently in the Daegok-ri dinosaur tracksite (Ulsan Metropolitan City Cultural Properties No. 13), Ulju-gun, Ulsan. Those tracks were reported as being the first in the Cretaceous deposits of Ulsan area. This tracksite is located relatively far from the Gyeongsangnam-do coastal area where pterosaur tracks were reported intensively, and there are stratigraphic differences. This study described the pterosaur tracks and trackway of the Daegok-ri tracksite and discussed the spatio-temporal distribution of the Cretaceous pterosaurs in the Korean Peninsula. The pterosaur footprints founded in Daegok-ri can be classified as Pteraichnus ichnosp., followed by the morphological characteristics. Based on the distribution of the pterosaur tracks on the Korean Peninsula including the Daegok-ri pterosaur tracks, it was estimated that the Cretaceous pterosaurs on the Korean Peninsula had survived approximately for 40 million years from 120 Ma (i.e., Early Cretaceous) to 80 Ma (i.e., Late Cretaceous) and inhabited in various sedimentary environments regardless of the basin size and the specific sedimentary environments. Furthermore, the occurrence of the sauropod, ornithopod, theropod, pterosaurs, birds, and crocodyliforms tracks/trackways in the Daegok-ri area represented that the Cretaceous vertebrate ichnodiversity was fairly high.

울산 울주군 대곡리 공룡 발자국 화석산지(울산광역시 문화재자료 제13호)에서 익룡 발자국이 새롭게 발견되었다. 이들은 울산 지역 백악기 퇴적층에서 처음 보고된 것이다. 이 발자국 화석산지는 익룡 발자국이 집중적으로 보고되었던 경상남도 해안 지역과 지리적으로 비교적 원거리에 위치하고, 층서적인 차이도 있다. 본 연구에서는 대곡리의 익룡 발자국 및 보행렬을 기재하였고, 백악기 동안 한반도에서 활동한 익룡의 시공간적 분포를 논의하였다. 대곡리에서 발견된 익룡 발자국은 형태적인 특징에 근거해 Pteraichnus ichnosp.로 분류할 수 있다. 대곡리 익룡 발자국을 포함하여 지금까지 알려진 한반도 익룡 발자국의 분포를 근거로 할 때, 한반도의 익룡들은 적어도 약 1억2천 만 년 전(120 Ma, 전기 백악기)부터 8천만 년 전(80 Ma, 후기 백악기)까지 약 4천만 년 동안 생존하였으며, 분지 규모나 특정 퇴적 환경에 관계없이 다양한 퇴적 환경에서 서식하였다. 그리고 대곡리 일대에서 용각류, 조각류, 수각류, 익룡, 새 그리고 악어류 발자국이 함께 산출되는 것은 이곳의 백악기 척추동물 생흔 다양성이 매우 높았다는 것을 의미한다.

Keywords

References

  1. Ambroggi, R. and Lapparent, A.F., 1954, Les empreintes de pas fossils du Maestrichtien d'Agadir. Notes du Service Gelogique du Maroc, 10, 43-57.
  2. Billon-Bruyat, J.P. and Mazin, J.M., 2003, The systematic problem of tetrapod ichnotaxa: thecase study of Pteraichnus Stokes, 1957 (Pterosauria, Pterodactyloidea). In Buffetaut, E., Mazin, J.M. (eds.), Evolution and palaeobiology of pterosaurs, vol. 217. Geological Society of London, pp. 315-324.
  3. Calvo, J.O. and Lockley, M.G., 2001, The first pterosaur track record in Gondwanaland. Cretaceous Research, 22, 585-590. https://doi.org/10.1006/cres.2001.0276
  4. Chough, S.K. and Sohn, Y.K., 2010, Tectonic and sedimentary evolution of a Cretaceous continental arcbackarc system in the Korean Peninsula: New view. Earth-Science Reviews, 101, 225-249. https://doi.org/10.1016/j.earscirev.2010.05.004
  5. Chun, S.S. and Chough, S.K., 1992, Tectonic history of Cretaceous sedimentary basins in the southwestern Korean Peninsula and Yellow Sea. In Chough, S.K. (ed.) Sedimentary basins in the Korean Peninsula and adjacent seas. Korean Sedimentology Research Group Special Publication, pp. 60-76.
  6. Delair, J.B., 1963, Notes on Purbeck fossil footprints, with descriptions of two hitherto unknown forms from Dorset, with notes on their stratigraphic distribution. Proceedings of the Dorset Natural History and Archaeological Society, 42, 293-322.
  7. Fuentes Vidarte, C. and Meijide Calvo, M., 1999, Primeras Huellas de Cocodrilo en el Weald de Cameros (Sria, Espana) Nueva Familia Crocodilopodidae: Nuevo icnogenero: Crocodylopodus Nueva icnoespecie: C. meijidei. Actas de las jornadas internacionales sobre paleontologia de dinosairios y su entorno. Sala de los infantes (Burgos, Espana). Collectivo Arqueologico-Paleontologico de Salas: 329-338.
  8. Ha, S., Yoo, J.S., Kim, K.S., Lim, H.S., Kang, S.H., Lockley, M.G., Lim, J.D., and Xing, L., 2016, New small pterosaur tracks from the Early Cretaceous in Jinju Formation, In. Abstract book of 2016 Annual Fall Joint Geology Meeting, 128 pp.
  9. Ha, S.W., Kim, J.M., Kim, K.S., Son, M.J., Choi, J.W., and Mun, S.G., 2013, Pterosaur trackways from the Cretaceous Jindong Formation of Jinju, Gyeongnam, Korea. In. Proceedings of the 2012 Fall Meeting of the Korean Earth Science Society, pp. 179-180.
  10. Hitchcock, E., 1858, Ichnology of New England, a report on the sandstones of the Connecticut Valley, Especially its fossil footmarks. William White, Boston, USA, 220 p.
  11. Hornung, J.J. and Reich, M., 2013, The first record of the Pterosaur ichnogenus Purbeckopus in the Late Berriasian (Early Cretaceous) of Northwest Germany. Ichnos, 20, 164-172. https://doi.org/10.1080/10420940.2013.845097
  12. Houck, K.J. and Lockley, M.G., 2006, Life in an active volcanic arc: Pterology and sedimentology of dinosaur track beds in the Jindong Formation (Cretaceous), Gyeongsang Basin, South Korea. Cretaceous Research, 27, 102-122. https://doi.org/10.1016/j.cretres.2005.10.004
  13. Huh, M., Lim, S.K., and Yang, S.Y., 1996, First discovery of pterosaur tracks from Asia. Journal of the Geological Society of Korea, 32, 526-528.
  14. Huh, M., Yang, S.Y., Seo, S.J., Kim, J.Y., Kim, K.S., Lim, J.D., and Kong, D.Y., 2009. Dinosaur fossils of Korea. Kungri Press, Daejeon, Korea, 297 p. (translated by Kim, J.Y. and Lee, S.Y., 2011, Kungri Press)
  15. Hwang, K.G., Huh, M., Lockley, M.G., Unwin, D.M., and Wright, J.L., 2002, New pterosaur tracks (Pteraichnidae) from the Late Cretaceous Uhangri Formation, southwestern Korea. Geological Magazine, 139, 421-435. https://doi.org/10.1017/S0016756802006647
  16. Jo, J.H., 2015, A perspective on the world heritage values of cultural landscape in the Daegokcheon Valley. Unpublished M.S. thesis, Korea National University of Education, Chungbuk, Korea, 113 p.
  17. Kang, H.C. and Paik, I.S., 2013, Review on the geological ages of the formations in the Gyeongsang Basin, Korea. Journal of the Geological Society of Korea, 49, 17-29.
  18. Kang, S.H., 2015, Paleoecology and vertebrate tracks from the Cretaceous Jinju Formation of Seopo-myeon, Sacheon City, Gyeongnam, Korea. Unpublished M.S. thesis, Chinju National University of Education, 50 p.
  19. Kim, C.B. and Kang, S.S., 2012. K-Ar Ages of Cretaceous fossil sites, Seoyuri, Hwasun, Southern Korea. Journal of the Korean Earth Science Society, 33, 618-626. https://doi.org/10.5467/JKESS.2012.33.7.618
  20. Kim, C.B., Huh, M., Cheong, C.S., Lockley, M.G., and Chang, H.W., 2003, Age of the pterosaur and webfooted bird tracks associated with dinosaur footprints from Korea. The Island Arc, 12, 125-131. https://doi.org/10.1046/j.1440-1738.2003.00385.x
  21. Kim, C.B., Kim, K.S., and Kim, N.H., 2014a, Geologic age of the Jinju Formation in the Hotandong, Jinju, In. Proceedings of the 2014 Spring Meeting of the Korean Earth Science Society, pp. 33.
  22. Kim, C.B., Kim, K.S., Kim, N.H., and Turek, A., 2014b, Geologic age and implication of the Cretaceous dinosaur track-bearing strata, Sanbukdong, Gunsan, In. Proceedings of the 2014 Fall Meeting and 50th founding anniversary of the Korean Earth Science Society, pp. 93.
  23. Kim, C.B., Kim, U.J., Park, M.S., Hwang, K.G., and Lee, K.W., 2017, SHRIMP U-Pb ages of dinosaur and bird footprints found in Cretaceous formation of Saok Island, Jeollanam-do, South Korea. Journal of the Korean Earth Science Society, 38, 141-149. https://doi.org/10.5467/JKESS.2017.38.2.141
  24. Kim, H.J. and Paik, I.S., 2017, Preliminary study on the new pterosaur footprints from the early Cretaceous Hasandong Formation on Hadong-gun, Gyeonsangnamdo, South Korea, In. Proc. Society of Vertebrate Paleontology 77th Annual Meeting, pp. 142.
  25. Kim, H.J., Hwang, K.G., Park, J.Y., and Huh, M., 2012, New pterosaur footprint fossils from the Korean Cretaceous Dinosaur Coast: Evidence of pterosaur diversity in the Cretaceous terrestrial ecosystems. In. The 11th Symposium on Mesozoic Terrestrial Ecosystems, 271-272.
  26. Kim, H.J., Paik, I.S., and Lim, J.D., 2014, Dinosaur trackbearing deposits at Petroglyphs of Bangudae Terrace in Daegokcheon stream, Ulju (National Treasure No. 285): Occurrences, paeoenvironments, and significance in natural history. Munhawjae Korean Journal of Cultural Studies, 47, 46-67.
  27. Kim, J.Y. and Lockley, M.G., 2016, First report of turtle tracks from the Lower Cretaceous of Korea. Cretaceous Research, 64, 1-6. https://doi.org/10.1016/j.cretres.2016.03.014
  28. Kim, J.Y., Kim, M.K., Oh, M.S., and Lee, C.Z., 2013, A new semi-palmate bird track, Gyeongsangornipes lockleyi ichnogen. et ichnosp. nov., and Koreanaornis from the Early Cretaceous Jindong Formation of Goseong County, southern coast of Korea. Ichnos, 20, 72-80. https://doi.org/10.1080/10420940.2013.787071
  29. Kim, J.Y., Kim, S.H., Kim, K.S., and Lockley, M.G., 2006, The oldest record of webbed bird and pterosaur tracks from South Korea (Cretaceous Haman Formation, Changseon and Sinsu islands): More evidence of high avian diversity in East Asia. Cretaceous Research, 27, 56-69. https://doi.org/10.1016/j.cretres.2005.10.005
  30. Kim, J.Y., Lockley, M.G., and Chun, H.Y., 2016, New dinosaur tracks from the Lower Cretaceous (Valanginian-Hauterivian) Saniri Formation of Yeongdong area, central Korea: Implications for quadrupedal ornithopod locomotion. Cretaceous Research, 61, 5-16. https://doi.org/10.1016/j.cretres.2015.12.023
  31. Kim, J.Y., Lockley, M.G., Kim, K.S., Seo, S.J., and Lim, J.D., 2012, Enigmatic giant pterosaur tracks and associated ichnofauna from the Cretaceous of Korea: Implication for the bipedal locomotion of pterosaurs. Ichnos, 19, 50-65. https://doi.org/10.1080/10420940.2011.625779
  32. Kim, K.S, Kim, J.M., Son, M.J., Ha, S.W., Choi, J.W., and Lim, J.D., 2013, Occurrence of pterosaur tracks in the Natural Monument No. 534 Pterosaur, Bird, and Dinosaur Tracks Fossil Site, Hotandong, Jinju, Korea: World richest pterosaur tracksite, In. Proceedings of the Geological Society of Korea, 342 p.
  33. Kim, K.S., Chung, C.H., Oh, K.H., Ha, S., Chae, Y.U., and Lim, H.S., 2017b, Vertebrate trace assemblage from the Cretaceous upper Haman Formation, Hyangchon area, Sacheon City, Gyeongnam, In. Proceeding of 2017 Spring Meeting of the Korean Earth Science Society, 64.
  34. Kim, K.S., Kang, S.H., Choi, Y.G., Kim, J.H., Bae, S.I., Shin, D.H., Yoo, J.S., and Kim, N.J., 2016a, A report on the attendance investigation in the construction site for housing of E-4 block, Jinju Innovation City. Korea Institute of Geoheritage, 41 p.
  35. Kim, K.S., Kim, C.B., Choi, Y.G., Kim, J.M., Ha, S.W., Mun, S.G., Kang, S.H., Choi, J.W., Son, M.J., Lee, H.M., and Kim, N.J., 2015, A report on the excavation of the fossil heritage in the Gasan General Industrial Complex, Gasan-ri, Ibanseong-myeon, Jinju, Gyeongnam. Korea Geoheritage Research Center, 190 p.
  36. Kim, K.S., Kim, C.B., Paik, I.S., Chung, C.H., Ahn, J.H., Seo, S.J., Kim, S.H., Kang, S.H., Choi, Y.G., Moon, S.K., Seo, H.S., Seol, W.K., Kim, Y.H., Kim, D.H., Ha, S.J., Yoo, J.S., Moon, H.W., Mun, S.C., Lee, S.H., Yoon, B.S., Park, H.D., Shin, D.H., Kim, J.H., Bae, S.I., Kim, N.J., 2016c, A report on the 3rd excavation of the fossil heritage at the Jinju Innovation City, Jinju. Gyeong Nam Development Corporation & Haedong Institute of Cultural Properties, 462 p.
  37. Kim, K.S., Kim, J.H., Bae, S.I., Shin, D.H., Yoo, J.S., and Lim, J.D., 2016b, A preliminary report on the vertebrate tracks and fossil plants from the attendance investigation of the construction site for housing of E-4 block, Jinju Innovation City, In. Proceedings of the 2016 Spring Meeting of the Korean Earth Science Society and Gyeongnam Goseong International Dinosaur Symposium, 131 p.
  38. Kim, K.S., Lim, J.D., and Ha, S., 2017c, Pterosaur tracks in the Daegok-ri, Ulju County, Ulsan City, In. Proceeding of 2017 Fall Meeting of the Korean Earth Science Society, 87 p.
  39. Kim, K.S., Lim, J.D., Kim, C.B., Kim, J.M., Ha, S.W., Son, M.J., and Choi, J.W., 2014a, A preliminary report on the dinosaur and pterosaur tracksite from Sanbukdong, Gunsan City, Jeonbuk, In. Proceedings of the 2014 Spring Meeting of the Korean Earth Science Society, 179-180.
  40. Kim, K.S., Lim, J.D., Lockley, M.G., Xing, L., and Choi, Y., 2017b, Korean trackway of a hopping, mammaliform trackmaker is first from the Cretaceous of Asia. Cretaceous Research, 74, 188-191. https://doi.org/10.1016/j.cretres.2017.02.005
  41. Kim, K.S., Lim, J.D., Lockley, M.G., Xing, L., Ha, S.J., Kim, C.B., Paik, I.S., Ahn, J.H., and Mun, S.C., 2018, First reports of a distinctive theropod track assemblage from the Jinju Formation (Lower Cretaceous) of Korea provides strong correlations with China. Cretaceous Research, 81, 26-35. https://doi.org/10.1016/j.cretres.2017.08.005
  42. Kim, K.S., Lim, J.D., Son, M.J., Kim, J.M., Ha, S.W., and Choi, J.W., 2014c, Pterosaur tracks from Jinseongmyeon, Jinju City, Gyeongsang nam-do, In. Proceedings of the 2014 Fall Meeting and 50th founding anniversary of the Korean Earth Science Society, 94 p.
  43. Kim, K.S., Lockley, M.G., Lim, J.D., Pinuela, L., Xing, L., and Moon, H.W., 2017a, First report of lacertiform (lizard) tracks from the Cretaceous of Asia. Cretaceous Research, 69, 62-70. https://doi.org/10.1016/j.cretres.2016.08.013
  44. Kim, K.S., Seo, S.J., Chung, C.H., Oh, K.H., Ha, S., Chae, Y.U., and Kim, N.J., 2017a, A summary report on the prospecting investigation of the natural properties in the construction site of Hyangchon 2 General Industrial Complex. Korea Institute of Geoheritage, 25 p.
  45. Kim, K.S., Seo, S.J., Kim, J.M., Ha, S.W., Choi, J.W., and Son, M.J., 2014b, A report of the collection investigation in the Sanbukdong dinosaur tracksite. Korea Geoheritage Research Center, 100 p.
  46. Kim, N.J., Kwon, Y.I., and Jin, M.S., 1971, Explanatory text of the geological map of Moryang Sheet. Geological Survey of Korea, 19 p.
  47. Kim, S.H., 2008, Stratigraphic diversity of bird and diminutive dinosaur tracks from the Cretaceous Haman Formation, South Gyeongsang Province of Korea: Implication for ichnostratigraphy. Unpublished Ph. D. dissertation, Korea National University of Education, Chungbuk, Korea, 141 p.
  48. Kong, D.Y. and Jung, S.H., 2018, A press release. http://www.nrich.go.kr/kor/boardView.do?menuIdx=283&bbscd=33&bbs_idx=40706 (June 5th 2018)
  49. Lee, D.W., 1999, Strike-slip fault tectonics and basin formation during the Cretaceous in the Korean Peninsula. Island Arc, 8, 218-231. https://doi.org/10.1046/j.1440-1738.1999.00233.x
  50. Lee, M.S. and Kang, P.C., 1964, Explanatory text of the geological map of Yangsan Sheet. Geological Survey of Korea, 24 p.
  51. Lee, Y.J. and Lee, I.K., 1972, Explanatory text of the geological map of Eon Yang Sheet. Geological Survey of Korea, 16 p.
  52. Lee, Y.N., Lee, H.J., L, J., and Kobayashi, Y., 2008, New pterosaur tracks from the Hasandong Formation (Lower Cretaceous) of Hadong County, South Korea. Cretaceous Research, 29, 345-353. https://doi.org/10.1016/j.cretres.2007.05.004
  53. Li, D., Xing, L., Lockley, M.G., Pinuela, L., Zhang, J., Dai, H., Kim, J.Y., Scott Persons IV, W., and Kong, D., 2015, A manus dominated pterosaur track assemblage from Gansu, China: Implications for behavior. Science Bulletin, 60, 264-272. https://doi.org/10.1007/s11434-014-0681-z
  54. Lim, J.D., Kong, D.Y., Kim, K.S., and Kim, T.H., 2009, Unusual quadrupedal ornithopod footprints and large manus imprint of pterosaur from the Early Cretaceous of Korea, In. 69th Annual Meeting Society of Vertebrate Paleontology and the 57th Symposium of Vertebrate Paleontology and Comparative Anatomy, 135 p.
  55. Lim, J.D., Kong, D.Y., Kim, T.H., Jeong, S.H., and Yoo, Y.W., 2012a, Geoheritage in the Daegokcheon Stream Petroglyphs district. National Research Institute of Cultural Heritage, 57 p.
  56. Lim, J.D., Kong, D.Y., Kim, T.H., Jeong, S.H., and Yoo, Y.W., 2012b, Geodiversity of Korea-Ulsan. National Research Institute of Cultural Heritage, 187 p.
  57. Lockley, M.G. and Meyer, C., 2004, Crocodylomorph trackways from the Jurassic to Early Cretaceous of North America and Europe: Implications for ichnotaxonomy. Ichnos, 11, 167-178. https://doi.org/10.1080/10420940490428832
  58. Lockley, M.G., 1999, Pterosaur and bird tracks from a new locality in the Late Cretaceous of Utah. In Gillette, D.D. (ed.), Vertebrate paleontology in Utah. Utah Geological Survey, Miscellaneous Publications, pp. 355-359, 99(1).
  59. Lockley, M.G., Huh, M., Kim, J.Y., Lim, J.D., and Kim, K.S., 2012, Recent advances in Korean vertebrate ichnology: The KCDC comes of age. Ichnos, 19, 1-5. https://doi.org/10.1080/10420940.2012.660400
  60. Lockley, M.G., Huh, M., Lim, S.K., Yang, S.Y., Chun, S.S., and Unwin, D.M., 1997, First report of pterosaur tracks from Asia, Chullanam province, Korea. Journal of the Paleontological Sociey of Korea. Special Publication, 2, 17-32.
  61. Masrour, M., de Ducla, M., Billon-Bruyat, J.-P., and Mazin, J.-M., 2018, Rediscovery of the Tagragra tracksite (Maastrichtian, Agadir, Morocco): Agadirichnus elegans Ambroggi and Lapparent 1954 is pterosaurian ichnotaxon. Ichnos, DOI:10.1080/10420940.2017.1386661.
  62. Masrour, M., Pascual-Arribas, C., de Ducla, M., Hernndez-Medrano, N., and Prez-Lorente, F., 2017, Anza palaeoichnological site. Late Cretaceous. Morocco. Part I. The first African pterosaur trackway (manus only). Journal of African Earth Science, 134, 766-775. https://doi.org/10.1016/j.jafrearsci.2017.07.004
  63. Mazin, J.M., Billon-Bruyat, J.P., Hantzpergue, P., and Lafaurie, G., 2003, Ichnological evidence for quadrupedal locomotion in pterodactyloid pterosaurs: Trackways from the Late Jurassic of Crayssac (Southwestern France). Geological Society, London, Special publication 217, 283-296. https://doi.org/10.1144/GSL.SP.2003.217.01.17
  64. McKeever, P., 2009, World heritage nomination-IUCN technical evaluation: Korean Cretaceous Dinosaur Coast (Republic of Korea), ID No. 1320. In IUCN evaluation of nominations of natural and mixed properties to the World Heritage List. WHC.09/33.COM/INF.8B2, pp. 12-22.
  65. Park, H.D., Kim, K.S., Lim, J.D., Kim, H.D., 2016, A preliminary study on crocodile tracks from the Cretaceous Jinju Formation of Jinju City, Gyeongnam. In Kim, K.S., Kim, D.H., and Lim, J.D. (eds.), 2016 Spring Meeting of the Korean Earth Science Society and Gyeongnam Goseong International Symposium, Gyeongnam Educational Welfare Center April 7-8th 2016, 143 p.
  66. Park, W.M., 2018, Upper Cretaceous vertebrate tracks and ichnodiversity from Saok and Song islands, Sinan County, Jeollanam-do, Korea. Unpublished M.S. thesis, Korea National University of Education, Chungbuk, Korea, 156 p.
  67. Park, W.M., Lockley, M.G., Kim, J.Y., and Kim, K.S., 2018, Anuran (frog) trackways from the Cretaceous of Korea. Cretaceous Research, 86, 135-148. https://doi.org/10.1016/j.cretres.2018.02.002
  68. Park, Y.D. and Yoon, H.D., 1968a, Explanatory text of the geological map of Ulsan Sheet. Geological Survey of Korea, 20 p.
  69. Park, Y.D. and Yoon, H.D., 1968b, Explanatory text of the geological map of Bangeujin Sheet. Geological Survey of Korea, 14 p.
  70. Pascual-Arribas, C., Canudo, J.I., Sanz Prez, E., Hernndez-Medrano, N., Castanera, D., and Barco, J.L., 2015, On the validity of Pteraichnus palacieisaenzi, Pascual Arribas and Sanz Prez 2000: New data from the Hurteles Formation. Palontologische Zeitschrift, 89, 459-483. https://doi.org/10.1007/s12542-014-0229-1
  71. Ryang, W.H., 2013, Characteristics of strike-slip basin formation and sedimentary fills and the Cretaceous small basins of the Korean Peninsula. Journal of the Geological Society of Korea, 49, 31-45.
  72. Ryu, C.K., Choi, M.K., Hong, S.H., Kim, S.H., and Kim, S.J., 2016, A report on the basic investigation research of the Cheonjeonri and Daegokri dinosaur tracksites, Ulsan. Korea Geological Environmental Institute and Ulsan Metropolitan City, 133 p.
  73. Ryu, I.C., Choi, S.G., and Wee, S.M., 2006, An inquiry into the formation and deformation of the Cretaceous Gyeongsang (Kyongsang) Basin, Southeastern Korea. Journal of the Korean Society of Economic and Environmental Geology, 39, 129-149.
  74. Seo, S.J., 1997, Lower Cretaceous dinosaur's-pterosaur's footprints from Haman Formation, Namhae, Kyongnam Korea. Research in Science Education, 23, 1-9.
  75. So, J.Y. and Kong, D.Y., 2018, A press release. http://www.nrich.go.kr/kor/boardView.do?menuIdx=283&bbscd=33&bbs_idx=40690(May 24th 2018)
  76. Stokes, W.L., 1957, Pterodactyl tracks from the Morrison Formation. Journal of Paleontology, 31, 952-954.
  77. Xing, L., Lockley, M.G., Pinuela, L., Zhang, J., Klein, H., Li, D., and Wang, F., 2013b, Pterosaur trackways from the Lower Cretaceous Jiaguan Formation (Barremian-Albian) of Qijiang, southwest China. Palaeogeography, Palaeoclimatology, Palaeoecology, 392, 177-185. https://doi.org/10.1016/j.palaeo.2013.09.003
  78. Xing, L.D., Lockley, M.G., Klein, H., Zhang, J.P., He, Q., Divay, J.D., Qi, L.Q., and Jia, C.K., 2013a, Dinosaur, bird and pterosaur footprints from the Lower Cretaceous of Wuerhe asphaltite area, Xinjiang, China, with notes on overlapping track relationships. Palaeoworld, 22, 42-51. https://doi.org/10.1016/j.palwor.2013.03.001
  79. Yang, S.Y., Choi, H.I., Seo, S.J., Kim, J.Y., and Kim, K.S., 2006, 2006 Report on the survey and research of fossil sites in southern coast (III). Cultural Heritage Administration of Korea, 149 p.
  80. Yang, S.Y., Choi, H.I., Seo, S.J., Kim, J.Y., and Kim, K.S., 2008, 2008 Report on the survey and research of fossil sites in southern coast (V). Cultural Heritage Administration of Korea, 133 p.
  81. Yonhap News Agency, 2018, Morphology and gait of major vertebrate tracks and trackways. http://img.yonhapnews.co.kr/photo/yna/YH/2018/06/05/PYH2018060517760001300_P4.jpgcid=PYH20180605177600013&from=search (June 5th 2018)