Acknowledgement
The authors are grateful to two anonymous reviewers for their invaluable comments, to Hwa-Jung Suh and Dong-Hyuk Lee for their help in the fieldwork, and to Yun Gyeong Choi for the assistance of laboratory work. This work was supported by the National Institute of Biological Resources under grant NIBR202005201 and the National Research Foundation of Korea under grant NRF-2020R111A3068464.
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